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Table 2 Marking behavioural units displayed by pampas deer males (Ozotoceros bezoarticus).

From: Reproductive biology of the pampas deer (Ozotoceros bezoarticus): a review

Behavioural unit

Behaviour in the male

Frontal marking

Brisk swinging head motions, either lateral or vertical, rubbing both antlers, simultaneously or alternately. Frequently performed with the tail raised, showing the perineal region.

Rubbing

A beating and twisting of the brush and small limbs, initially in the stripping of the velvet and consequently, although perhaps incidentally, in the polishing of the antlers

Antler trashing

The chafing of the basal antler parts against the trunks of small trees

Pre-orbital marking

Stands with the neck stretched, pressing the gland against an object, with short and energetic up and down movements

Front leg marking

Performed with the neck slightly below the shoulder height, scratching the ground with the front hooves, frequently alternating the hooves